Chuck , Sarah and Casey are shocked to discover that Chuck’s girlfriend Jill (guest star Jordana Brewster) is a Fulcrum agent in search of the Intersect. In the wake of his disbelief, Chuck is asked to use his relationship with Jill to get to an agent called Leader. He willingly agrees to bring down the woman who betrayed him twice. Meanwhile, Devon's parents are coming for Thanksgiving and Ellie is determined to have the perfect Thanksgiving dinner. To make sure everything goes smoothly, she tells Morgan that he cannot come to dinner this year. Without any holiday plans to fall back on, Morgan is recruited by Big Mike to stand guard at the Buy More with Lester and Jeff.

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